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To infer the age of ancient genomes, we take advantage of the shared history of Neanderthal gene flow into non-Africans that occurred around 50,000 y ago and measure the amount of “missing evolution” in terms of recombination breakpoints in the ancient genome compared with present-day samples.The study of human evolution has been revolutionized by inferences from ancient DNA analyses.Key to these studies is the reliable estimation of the age of ancient specimens. The first direct radiocarbon dating of five human fossils from Mlade is now available. Although generally accepted as modern Homo sapiens, features of the cranial morphology have prompted speculation about anatomical links to the preceding Neanderthals. The human remains from the Mlade caves in the Czech Republic feature strongly in discussions of the transition from Neanderthals to early modern humans in Europe.

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