(It went poorly at first, and Bonecrusher offering to offline the two Autobots did not help.) While building the tower, they explained to Megatron (at gunpoint) that they were still loyal to him, and once the tower was built, they would turn it over to him. Present at the Battle of Autobot City, Bonecrusher helped to breach the Autobot defenses as part of Devastator, who withstood an Autobot attack.
After all was said and done, the Autobots put Bonecrusher and the rest of the Constructicons to work, restoring the island to its former natural splendor.Bonecrusher and the Constructicons were stationed on Earth in 2011, under the command of Sixshot.They attempted to prevent Ultra Magnus from sending reinforcements to Cybertron to help the Autobots fend off Galvatron's siege on Vector Sigma.Megatron, wanting their constructicons skills on his side, reprogrammed them to serve him using the Robosmasher.Megatron built Bonecrusher, who, with his fellow Constructicons, would attempt to deal with Teletraan I, only to be thwarted by the Dinobots.Bonecrusher and his squad were ganging up on the Trainbots until Metroplex arrived and shooed them off with his superior firepower.
Having failed to stop the Autobot reinforcements from departing, the Constructicons followed them to Cybertron.
After Devastator sealed the Decepticons in a tunnel and helped rebuild some of the Autobot base, Megatron then revealed he knew of the Autobot plan to take control of the Constructicons, and deactivated the disks.
After a fight for control, Bonecrusher, as Devastator, goes berserk.
He loves to show off his destructive talents for his fellow Decepticons.
Deep down, his destruction is really a perfectionism that cannot tolerate anything that he views as flawed, which includes the Autobots.
(And who should be thrown out of the shuttle.) Though Rumble and Frenzy prevented Devastator from proving his superiority, Bonecrusher and the other Constructicons made it back to Cybertron and were present for Starscream's coronation as Decepticon leader.